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The film follows the Turtle brothers as they work to earn the love of New York City while facing down an army of mutants.
Fifteen years ago, a Techno Cosmic Research Institute strike force is ordered by executive Cynthia Utrom to capture rogue scientist Baxter Stockman, who stole company secrets and technology, and any research. Desiring a family, Stockman creates a mutagen to create his own mutant family, starting with a housefly maggot. Before he can create more mutagen, the strike force arrives. While it is dealt with by the fly mutant, an explosion in the fight destroys the lab and most of the research while killing Stockman. Additional TCRI forces arrive and collect the remainder of the surviving research, but forget the mutant and the last canister of mutagen, the latter of which falls into the sewers below.
In the present, in New York City, four turtles (Michaelangelo, Leonardo, Raphael, and Donatello) have been raised by their adoptive father Splinter. While they are adept in the arts of ninjitsu, they are instructed to only leave for the surface world to get supplies for their home in the sewers.
During these runs, they occasionally go and see attractions like outdoor movies, concerts, and sports, which makes them want to be on the surface more. When Splinter discovers what they have been doing, he punishes them, explaining their backstory: the mutagen from Stockman's house fell on the four baby turtles, who were found by Splinter (then a rat that was cynical and untrusting of the surface world, specifically of humans). While handling the turtles, Splinter also comes into contact with the mutagen, mutating all of them to have more human-like features. As the Turtles grew older and became more curious about the surface, he decided to take them to the surface where they are all nearly killed. This leads Splinter and the Turtles to all learn martial arts for their protection.
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